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Cyperus
Cyperus resembles Kyllinga,
but Cyperus has > 5 spikelet scales, whereas Kyllinga has 1-3
scales.
Sometimes classified as "Cyperus pedunculatus,"
Remirea maritima is a small-statured maritime species on beach dunes.
See

1. Plants with numerous (>30) inflorescence rays
and no (or nearly no) bracts… (Cyperus papyrus,
another garden species, may have > 30 rays but is much larger to > 2 m tall vs. < 1 m tall)
1. Plants otherwise…2
2. Stigmas 3; achenes 3-angled…8
2. Stigmas 2; achenes 2-angled…3
Species with 2 styles and
flattened achenes:
3. Spikelets crowded densely into cylindric
spikes… (a large invasive exotic species not yet
reported in our counties but encountered nearby)
3. Spikes not densely crowded (rachis visible
between the spikelets, or if the rachis not visible, the plant then with a
total of < 100 spikelets vs. too many to count in C. alopecuroides)…4
4. Scales awned, the margins white…
4. Scales not awned (may have a tiny cusp), the
margins not white…5
5. Spikelets 3-5 mm wide
(rare)
5. Spikelets to 3 mm wide...6
6. Achene much longer than wide; spikelets
1.3-1.6(2) mm wide… (very common weed and highly
variable, the spikelets variable in length)
6. Achene ellipsoid or ovate; spikelets > 2 mm
wide…7
7. Mature achene black or brown, the surface
covered with elongate cells…
7. Mature achene brown, the surface covered with
square cells…
Species with 3 styles and
triangular achenes:
8. Culm scabrous…16 (See also C. ligularis
with a slightly roughened culm)
8. Culm smooth…9
9. Stems hollow, to 2 m tall; spikelets
5 cm long; bract appearing to be an extension of the culm making the
spikelets appear to be lateral (then resembling
Schoenoplectus, which differs by having > 1 achene per
much-shorter spikelet), or the spikelets in an umbellate arrangement at
the top of the culm; leaves and culms septate… (rare)
9. Plants otherwise…10
10. Plants > 1 m tall, having > 40 rays, the culm >
1.5 cm in diameter… (C.
papyrus, an escaped horticultural
species)
10. Plants not with the above combination of
characters…11
11. Plants with >17 broad flat bracts (these often
sticking straight out horizontally or nearly so) …
(Umbrella Sedge, an escaped horticultural species)
11. Plants otherwise…12
12. Achenes with spongy white base…
(looks like C. surinamensis but with smooth stem and with the
characteristic white achene base)
12. Achenes without white spongy base…13
13.
Spikelets in elongate cylindric spikes, the rachis breaking apart
and having
the segments remaining with the achenes. (A large common
eye-catching species of stagnant ditches and shores.
Similar,
C. strigosus has spikes with a bottlebrush appearance resembling
C. odoratus, but C. odoratus has round vs. flat spikelets
0.8-1.9 vs. 0.6-0.8 mm wide.)
13. Rachis not cylindric, not with corky wings, not
breaking apart with the achenes14
14. Spikelets in dense heads (with distinct globose
or cylindric shape)…15 (Note: If plant has a single sessile head > 3 cm
diam. made of numerous long spikelets, consider anomalous specimen of
C. planifolius, a seashore species appearing below)
14. Spikelets loosely arranged (the rachis showing
between the bases, or the bases crowded or touching but the clusters
irregular in outline)…21
15. Spikelet clusters globose…17 (See also
Cyperus elegans which has septate viscid leaves)
15. Spikelet clusters cylindric…18
16. Stems retrorsely scabrous…
(C. distinctus has similar appearance but smooth stems and
achenes with white spongy bases)
16. Stems antrorsely scabrous… (C.
distinctus has similar appearance but smooth stems and achenes with
white spongy bases)
17. Spikelet scales and achenes 0.6-0.8 mm long… (resembles
but has achenes
0.6-0.8 mm (vs. achenes 2.2-2.8 mm) and without white covering)
17. Spikelet scales 2.3-3.6 mm long and achenes 2-2.4
mm long…
18. Spikelets 2.2-4 mm long; leaves 1.5-4 mm wide… (spikelets remain greenish, species of dry sandy habitats)
18. Spikelets 3-30 mm long (if spikelets < 4 mm
the leaves > 5 mm wide)…19
19. Leaves 1-4 mm wide… (Spikes
with a bottlebrush appearance resembling C. odoratus but C.
odoratus with a round vs. flat spikelet 0.8-1.9 vs. 0.6-0.8 mm
wide) (Cyperus rotundus has tubers and similarly
narrow leaves but reddish spikelets only rarely exceeding 4 mm long vs.
usually well over 5 mm long)
19. Leaves 5-13 mm wide…20
20. Spikelets 3-7 mm long; spikes usually < 1.5 cm
in diameter… (spikelets becoming brownish vs. remaining
green in C. retorsus)
20. Spikelets 6-22 mm long; spikes > 1.5 cm in
diameter. (achene 1.3-2 mm)
21. Inflorescences
vaguely paniculate with the second-order branches having a feathery-shaggy
appearance covered for most of their length with loosely arranged
spikelets…
21. Not as above…22
22. Spikelets red or with
reddish markings…23
22. Spikelets green or
yellowish or golden-brown…24
23. Scales white with two
red spots…
23. Scales reddish
throughout (except for margins and midrib)… (This and
C. esculentus are our only two tuber-producing species. Both are
common aggressive weeds)
24. Leaf bladeless (sheath
only)… (Stem strongly triangular. Spikelets 1-1.6 mm wide)
24. Plants with leaf
blades…25
25. Rachis long, visible
between the spikelets, these spaced out along the rachis…26
25. Rachis short, not
visible, the spikelets crowded basally in digitate arrangement…27
26. Plants with tubers; achene 1.3-1.5 mm long;
(1.2)1.4-2(3) mm wide...
26. Plants otherwise...29
27. Plants rhizomatous; spikelets per cluster
(1)2-4(6)…
27. Plants cespitose; spikelets per cluster
usually 3-20…28
28. Sheath septate, viscous (sticky); achene > 1.4 mm
long; spikelets per cluster 3-20(30)…
28. Sheath not septate, not viscous; achene
1.2-1.4 mm long; spikelets per cluster (1)3-12(16)…
29. Spikelet 5-30 mm long, 0.6-0.9 mm wide...
29. Spikelet 5-8 mm long, 1.5-2 mm wide...
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